I wouldn't be surprised Alan looking at his website that a lot of links were built naturally by people linking to his site as it looks like a good resource.
I've got a website were I built some links to gain a good position and left it, over a few months that exposure built tons of new links without me knowing from all over the web.
Now if those little sites have been hit, lets say a fair amount had been wiped out then I can see how rankings could be lost.